
Flowering shrub
Inkberry
Ilex glabra
Inkberry (Ilex glabra) is a shrub in the Aquifoliaceae family, native in the lower 48 states, blooming mostly April–June across the U.S.
Inkberry is a popular flowering shrub. It's on our flower lists for 31 states, and across the U.S. it's most often seen in bloom in April–June.
*iNaturalist research-grade observations, as of October 23, 2026. Our state lists also include states where people grow it in gardens, so they can outnumber USDA's wild-plant records. Planted garden shrubs are mostly excluded from research grade.
On our lists for: Alabama, Arkansas, California, Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Kansas, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Oklahoma, Oregon, Pennsylvania, Rhode Island, South Carolina, Tennessee, Vermont, Virginia, West Virginia
Quick look
Ilex glabra, also known as Appalachian tea, evergreen winterberry, Canadian winterberry, gallberry, inkberry, dye-leaves and houx galbre, is a species of evergreen holly native to the coastal plain of eastern North America, from coastal Nova Scotia to Florida and west to Louisiana where it is most commonly found in sandy woods and peripheries of swamps and bogs.
- It belongs to the Aquifoliaceae family. iNaturalist
- USDA PLANTS describes it as a shrub with a perennial life cycle. USDA PLANTS
- USDA PLANTS lists it as native in the lower 48 states and records it in 20 states. USDA PLANTS
- In Florida, most iNaturalist sightings of it in flower come in April–May. iNaturalist
- In New Jersey, most iNaturalist sightings of it in flower come in June. iNaturalist
- USDA PLANTS lists its flower color as white. USDA PLANTS
- USDA PLANTS gives its bloom period as spring. USDA PLANTS
- USDA PLANTS lists a mature height of about 8 ft. USDA PLANTS
- iNaturalist has 6,057 research-grade sightings of it in Florida, ranking #25 among the 151 flowers on our Florida list. iNaturalist
- iNaturalist has 1,439 research-grade sightings of it in New Jersey, ranking #62 among the 186 flowers on our New Jersey list. iNaturalist
- iNaturalist has 785 research-grade sightings of it in North Carolina, ranking #109 among the 192 flowers on our North Carolina list. iNaturalist
When does it bloom?
Bloom time in the 31 states whose Bloomipedia list includes it
| State | Type | Peak bloom | Sightings* |
|---|---|---|---|
| Florida | 🌷 Garden | April–May | 6,057 |
| New Jersey | 🌷 Garden | June | 1,439 |
| North Carolina | 🌷 Garden | few records (7) | 785 |
| Massachusetts | 🌷 Garden | few records (17) | 731 |
| Mississippi | 🌷 Garden | April–May | 429 |
| Alabama | 🌷 Garden | few records (19) | 401 |
| Georgia | 🌷 Garden | few records (11) | 344 |
| New York | 🌷 Garden | few records (2) | 342 |
| South Carolina | 🌷 Garden | few records (8) | 236 |
| Virginia | 🌷 Garden | few records (3) | 148 |
| Rhode Island | 🌷 Garden | few records (3) | 120 |
| Louisiana | 🌷 Garden | few records (1) | 45 |
| Maryland | 🌷 Garden | few records (1) | 45 |
| Pennsylvania | 🌷 Garden | few records (4) | 44 |
| Connecticut | 🌷 Garden | no flowering records | 27 |
| Delaware | 🌷 Garden | no flowering records | 16 |
| Maine | 🌷 Garden | few records (2) | 6 |
| Tennessee | 🌷 Garden | no flowering records | 6 |
| New Hampshire | 🌷 Garden | no flowering records | 2 |
| West Virginia | 🌷 Garden | no flowering records | 1 |
| Arkansas | 🌷 Garden | no flowering records | 0 |
| California | 🌷 Garden | no flowering records | 0 |
| Illinois | 🌷 Garden | no flowering records | 0 |
| Indiana | 🌷 Garden | no flowering records | 0 |
| Kansas | 🌷 Garden | no flowering records | 0 |
| Kentucky | 🌷 Garden | no flowering records | 0 |
| Missouri | 🌷 Garden | no flowering records | 0 |
| Ohio | 🌷 Garden | no flowering records | 0 |
| Oklahoma | 🌷 Garden | no flowering records | 0 |
| Oregon | 🌷 Garden | no flowering records | 0 |
| Vermont | 🌷 Garden | no flowering records | 0 |
Conservation status
- Federal (U.S. ESA)
- Not listed No U.S. Endangered Species Act listing in the USFWS ECOS database. USFWS ECOS
- NatureServe global
- G5 Secure worldwide. NatureServe
- State rare plant lists
- Not listed Not on the official rare plant list in 17 of our 31 states.
See each state
- Florida
- Not listed Not on Florida Regulated Plant Index (Fla. Admin. Code 5B-40.0055). NatureServe: S5, secure in Florida. FAC 5B-40.0055
- New Jersey
- Not listed Not on New Jersey Endangered Plant Species List (N.J.A.C. 7:5C-5.1). NatureServe: S5, secure in New Jersey. N.J.A.C. 7:5C-5.1
- North Carolina
- Not listed Not on North Carolina Protected Plant Species List (02 NCAC 48F .0301). NatureServe: S5, secure in North Carolina. 02 NCAC 48F .0301
- Massachusetts
- Not listed Not on Massachusetts Endangered Species Act list (321 CMR 10.90). NatureServe: SNR, not yet ranked in Massachusetts. MESA list
- Georgia
- Not listed Not on Georgia protected plants (Ga. Comp. R. & Regs. 391-4-10-.09; a 2026 update is proposed). NatureServe: SNR, not yet ranked in Georgia. Ga. R. 391-4-10-.09
- Virginia
- Not listed Not on Virginia endangered and threatened plant and insect species (2VAC5-320-10). NatureServe: S5, secure in Virginia. 2VAC5-320-10
- Rhode Island
- Not listed Not on Rhode Island Rare Plants 2016 (RI Natural History Survey, from the RI Wildlife Action Plan; informational, Rhode Island has no state plant-protection listing law). NatureServe: SNR, not yet ranked in Rhode Island. RI rare plants 2016
- Maryland
- Not listed Not on Maryland threatened and endangered plants (COMAR 08.03.08.05, .06 and .08). NatureServe: SNR, not yet ranked in Maryland. COMAR 08.03.08
- Delaware
- Not listed Not on Rare and Uncommon Plants of Delaware (DNREC, 2024; informational, Delaware has no state plant-protection list). NatureServe: S5, secure in Delaware. DE rare plants 2024
- Tennessee
- Not listed Not on Tennessee endangered plants (Tenn. Comp. R. & Regs. 0400-06-02-.04, Rare Plant Protection and Conservation Act). NatureServe has no Tennessee rank for it. TN rule 0400-06-02-.04
- California
- Not listed Not on California state-listed endangered, threatened and rare plants (14 CCR § 670.2), per CDFW's CNDDB list (July 2026). NatureServe has no California rank for it. CNDDB listed plants (July 2026)
- Illinois
- Not listed Not on Illinois List of Endangered and Threatened Flora (17 Ill. Adm. Code 1050), per the Endangered Species Protection Board's 2025 checklist. NatureServe has no Illinois rank for it. IL E&T checklist 2025
- Kentucky
- Not listed Not on Kentucky endangered and threatened plant lists (400 KAR 3:040, eff. 2026). NatureServe has no Kentucky rank for it. 400 KAR 3:040
- Missouri
- Not listed Not on Missouri Wildlife Code endangered species (3 CSR 10-4.111), plants. NatureServe has no Missouri rank for it. 3 CSR 10-4.111
- Ohio
- Not listed Not on Ohio endangered and threatened plants (Ohio Adm. Code 1501:18-1-03, eff. Dec. 26, 2024). NatureServe has no Ohio rank for it. OAC 1501:18-1-03
- Oregon
- Not listed Not on Oregon endangered and threatened plants (OAR 603-073-0070). NatureServe has no Oregon rank for it. OAR 603-073-0070
- Vermont
- Not listed Not on Endangered and Threatened Plants of Vermont (10 V.S.A. chap. 123; list of Feb 10, 2022). NatureServe has no Vermont rank for it. Vermont E&T list
- Mississippi
- No list Mississippi has no state law listing endangered or threatened plants, so we show NatureServe's Mississippi ranks instead. NatureServe: SNR, not yet ranked in Mississippi.
- Alabama
- No list Alabama has no state law listing endangered or threatened plants, so we show NatureServe's Alabama ranks instead. NatureServe: SNR, not yet ranked in Alabama.
- New York
- Exploitably vulnerable On New York protected native plants (6 NYCRR § 193.3): exploitably vulnerable (protected from picking without the landowner's permission). NatureServe: S4, apparently secure in New York. 6 NYCRR § 193.3
- South Carolina
- No list South Carolina has no state law listing endangered or threatened plants, so we show NatureServe's South Carolina ranks instead. NatureServe: S5, secure in South Carolina.
- Louisiana
- No list Louisiana has no state list of endangered or threatened plants separate from the federal list, so we show federal status and NatureServe's Louisiana ranks. NatureServe: SNR, not yet ranked in Louisiana.
- Pennsylvania
- Pennsylvania extirpated On Pennsylvania classified native plants (17 Pa. Code Ch. 45, Subchapter B, as amended 2023): pennsylvania extirpated. NatureServe: SX, presumed extirpated in Pennsylvania. 17 Pa. Code Ch. 45
- Connecticut
- State threatened On Connecticut Endangered, Threatened and Special Concern Plants (RCSA 26-306; 2015 list): state threatened. NatureServe: S2, imperiled in Connecticut. CT E/T/SC list
- Maine
- State endangered On Maine Rare Plant List (Maine Natural Areas Program, 2021). E and T are Maine's official endangered and threatened plants (12 M.R.S.); SC and PE are tracking categories: state endangered. NatureServe: S1, critically imperiled in Maine. Maine rare plant list
- New Hampshire
- State endangered On New Hampshire Official Rare Plants List (Ncr 312, eff. Jan 1, 2020): state endangered. NatureServe: SH, possibly extirpated (known only from historical records) in New Hampshire. NH Ncr 312
- West Virginia
- No list West Virginia has no state law listing endangered or threatened plants (only federally listed species are protected), so we show NatureServe's West Virginia ranks instead. NatureServe has no West Virginia rank for it.
- Arkansas
- No list Arkansas has no state law listing endangered or threatened plants, so we show NatureServe's Arkansas ranks instead. NatureServe: SNR, not yet ranked in Arkansas.
- Indiana
- No list Indiana has no state law listing endangered or threatened plants (the DNR keeps a rare plant list, but it isn't regulatory), so we show NatureServe's Indiana ranks instead. NatureServe has no Indiana rank for it.
- Kansas
- No list Kansas's endangered and threatened list (K.A.R. 115-15-1) covers animals only, so we show NatureServe's Kansas ranks for plants. NatureServe has no Kansas rank for it.
- Oklahoma
- No list Oklahoma has no state law listing endangered or threatened plants, so we show NatureServe's Oklahoma ranks instead. NatureServe has no Oklahoma rank for it.
- Rare elsewhere
- Other states where NatureServe ranks it rare or missing: TX (S1). (S1 critically imperiled, S2 imperiled, SH possibly extirpated, SX presumed extirpated.)
Checked October 23, 2026. Rankings and lists can change, so follow the links for the latest.
Please don't pick or dig this flower: it's on New York's rare plant list (exploitably vulnerable); it's on Pennsylvania's rare plant list (pennsylvania extirpated); it's on Connecticut's rare plant list (state threatened); it's on Maine's rare plant list (state endangered); it's on New Hampshire's rare plant list (state endangered); NatureServe ranks it S2 in Connecticut and S1 in Maine. 6 NYCRR § 193.3 17 Pa. Code Ch. 45 CT E/T/SC list Maine rare plant list NH Ncr 312 NatureServe
Can you eat it?
NC State Extension lists it as poisonous to humans (severity: low). Symptoms they list: “Minor toxicity. Ingestion may cause vomiting, diarrhea, or other illness in humans.” They also note: “Use caution. Human ingestion of berries can cause minor toxic reaction.” NC State Extension
It isn't on the ASPCA plant list, and NC State Extension doesn't flag it for pets. For context only: ASPCA lists a related species, Ilex aquifolium (“English Holly”), as toxic to dogs and cats. That doesn't tell us about this species, so keep pets away to be safe. ASPCA
🌿 Never eat a wild plant unless you're certain of its ID; this isn't medical advice. Worried about a person? Call Poison Control at 1-800-222-1222. Worried about a pet? Call your vet.
Where does it grow?
USDA PLANTS records inkberry in 20 states.
recorded by USDA PLANTS on our state lists not recorded
See the full list of states
Alabama, Arkansas, Connecticut, Delaware, Florida, Georgia, Louisiana, Maine, Maryland, Massachusetts, Mississippi, New Hampshire, New Jersey, New York, North Carolina, Pennsylvania, Rhode Island, South Carolina, Texas, Virginia.
Florida counties with USDA records: Alachua, Baker, Bay, Bradford, Brevard, Broward, Calhoun, Charlotte, Citrus, Clay, Collier, Columbia, DeSoto, Dixie, Duval, Escambia, Flagler, Franklin, Gadsden, Gilchrist, Glades, Gulf, Hamilton, Hardee, Hendry, Hernando, Highlands, Hillsborough, Holmes, Indian River, Jackson, Jefferson, Lafayette, Lake, Lee, Leon, Levy, Liberty, Madison, Manatee, Marion, Martin, Miami-Dade, Monroe, Nassau, Okaloosa, Okeechobee, Orange, Osceola, Palm Beach, Pasco, Pinellas, Polk, Putnam, Santa Rosa, Sarasota, Seminole, St. Johns, St. Lucie, Sumter, Suwannee, Taylor, Union, Volusia, Wakulla, Walton, Washington.
New Jersey counties with USDA records: Atlantic, Burlington, Camden, Cape May, Cumberland, Gloucester, Hudson, Mercer, Middlesex, Monmouth, Morris, Ocean, Salem.
North Carolina counties with USDA records: Beaufort, Bertie, Bladen, Brunswick, Carteret, Chatham, Chowan, Columbus, Craven, Cumberland, Dare, Duplin, Durham, Edgecombe, Gates, Greene, Halifax, Harnett, Hertford, Hoke, Hyde, Johnston, Jones, Lee, Martin, Moore, Nash, New Hanover, Northampton, Onslow, Orange, Pamlico, Pasquotank, Pender, Pitt, Richmond, Robeson, Sampson, Scotland, Tyrrell, Vance, Wake, Wayne, Wilson.
Growing conditions
USDA PLANTS gives Inkberry a high shade tolerance, low drought tolerance and high moisture use. It is rated for soil pH 4.5 to 7.0 and winter lows down to -13 °F.
- Shade tolerance
- High
- Drought tolerance
- Low
- Moisture use
- High
- Soil textures it is adapted to
- coarse, medium, fine
- Soil pH range
- 4.5 to 7.0
- Minimum temperature tolerated
- -13 °F
- Frost-free days needed (minimum)
- 165
- Annual precipitation range
- 32 to 60 inches
- Minimum root depth
- 16 inches
- Salinity tolerance
- Medium
- Fertility requirement
- Medium
- Growth rate
- Slow
- Lifespan
- Long
- Active growth period
- Summer and Fall
- Fruit or seed
- Black · Summer to Fall
- Can be propagated by
- bare root, container, seed
- Commercial availability
- Routinely Available
From the USDA PLANTS characteristics data for ILGL. USDA ratings are general guidance for the species, not a guarantee for your garden.
Names, range and relatives
Native status by region (USDA PLANTS): Canada: Native; Lower 48 states: Native.
County records: USDA PLANTS lists it in 271 counties across 16 states; the most are in Florida (67), North Carolina (44), Georgia (39).
Other Ilex on Bloomipedia (4): American holly, Yaupon holly, Winterberry, Japanese holly.
More Florida flowers
Sources
- USDA, NRCS. PLANTS Database, profile ILGL.
- iNaturalist, Ilex glabra: taxonomy, research-grade observation counts and “Flowering” annotations, retrieved October 23, 2026.
- Wikipedia, Ilex glabra (lead quoted under CC BY-SA 4.0).
- State rare and invasive plant lists (50):
Show all 50 state lists
- Florida: Florida Regulated Plant Index (Fla. Admin. Code 5B-40.0055).
- Florida: Florida Noxious Weed List (Fla. Admin. Code 5B-57.007).
- New Jersey: New Jersey Endangered Plant Species List (N.J.A.C. 7:5C-5.1).
- New Jersey: New Jersey prohibited invasive plant species (Invasive Species Management Act; propagation banned from Jan 1, 2027, sale from Jan 1, 2030).
- North Carolina: North Carolina Protected Plant Species List (02 NCAC 48F .0301).
- North Carolina: North Carolina Noxious Weed List (02 NCAC 48A .1702) (the list names any non-native Lythrum; we apply it to purple loosestrife (Lythrum salicaria), which isn't native).
- Massachusetts: Massachusetts Endangered Species Act list (321 CMR 10.90).
- Massachusetts: Massachusetts Prohibited Plant List (MDAR) (Text from MDAR's 2023 scientific-name PDF (mass.gov blocks automated downloads); 2024-2026 additions listed on the mass.gov page: Alnus glutinosa, Pyrus calleryana, Miscanthus sinensis, Aralia elata, Rhodotypos scandens, Symplocos paniculata.).
- Mississippi: Mississippi Noxious Weed List (2 Miss. Code R. 1-3-01-136.01).
- Alabama: Alabama noxious weeds (Ala. Admin. Code r. 80-10-14-.04).
- Georgia: Georgia protected plants (Ga. Comp. R. & Regs. 391-4-10-.09; a 2026 update is proposed).
- New York: New York protected native plants (6 NYCRR § 193.3).
- New York: New York prohibited and regulated invasive species (6 NYCRR Part 575).
- South Carolina: South Carolina State Plant Pest List, plants (S.C. Code Regs. 27-135; Clemson Dept. of Plant Industry).
- Virginia: Virginia endangered and threatened plant and insect species (2VAC5-320-10).
- Virginia: Virginia noxious weeds (2VAC5-317-20, Tier 1, 2 and 3).
- Rhode Island: Rhode Island Rare Plants 2016 (RI Natural History Survey, from the RI Wildlife Action Plan; informational, Rhode Island has no state plant-protection listing law).
- Maryland: Maryland threatened and endangered plants (COMAR 08.03.08.05, .06 and .08).
- Maryland: Maryland Prohibited Invasive Plant List (COMAR 15.06.04.06; MDA, updated July 2026).
- Pennsylvania: Pennsylvania classified native plants (17 Pa. Code Ch. 45, Subchapter B, as amended 2023).
- Pennsylvania: Pennsylvania Noxious Weed List (3 Pa.C.S. § 1519, PA Dept. of Agriculture) (PDA's page spells garlic mustard “Allaria petiolata”; we match it as Alliaria petiolata).
- Connecticut: Connecticut Endangered, Threatened and Special Concern Plants (RCSA 26-306; 2015 list).
- Connecticut: Connecticut prohibited invasive plants (Conn. Gen. Stat. § 22a-381d).
- Delaware: Rare and Uncommon Plants of Delaware (DNREC, 2024; informational, Delaware has no state plant-protection list).
- Delaware: Delaware Invasive Plant List (3 Del. Admin. Code § 806-3.0).
- Maine: Maine Rare Plant List (Maine Natural Areas Program, 2021). E and T are Maine's official endangered and threatened plants (12 M.R.S.); SC and PE are tracking categories.
- Maine: Maine Do Not Sell invasive plant list (01-001 C.M.R. ch. 273).
- Tennessee: Tennessee endangered plants (Tenn. Comp. R. & Regs. 0400-06-02-.04, Rare Plant Protection and Conservation Act).
- Tennessee: Tennessee Pest Plant List (Tenn. Comp. R. & Regs. 0080-06-24-.02).
- New Hampshire: New Hampshire Official Rare Plants List (Ncr 312, eff. Jan 1, 2020).
- New Hampshire: NH Prohibited Invasive Species List (Agr 3800, Table 3800.1).
- West Virginia: West Virginia noxious weeds (W. Va. Code R. § 61-14A-5) (the rule abbreviates some names (e.g. “P. cuspidatum”); we expand them to match).
- Arkansas: Arkansas State Plant Board noxious weeds and prohibited plants (2 CAR § 50-101).
- California: California state-listed endangered, threatened and rare plants (14 CCR § 670.2), per CDFW's CNDDB list (July 2026).
- California: California noxious weed species (3 CCR § 4500).
- Illinois: Illinois List of Endangered and Threatened Flora (17 Ill. Adm. Code 1050), per the Endangered Species Protection Board's 2025 checklist.
- Illinois: Illinois Official List of Exotic Weeds (17 Ill. Adm. Code 1100.30, Illinois Exotic Weed Act).
- Indiana: Indiana prohibited invasive terrestrial plants (312 IAC 18-3-25).
- Kansas: Kansas noxious weeds (K.A.R. 4-8-44, as amended 2026).
- Kentucky: Kentucky endangered and threatened plant lists (400 KAR 3:040, eff. 2026).
- Kentucky: Kentucky noxious weeds and invasive plants on state rights-of-way (KRS 176.051; species list from the Kentucky Transportation Cabinet).
- Missouri: Missouri Wildlife Code endangered species (3 CSR 10-4.111), plants.
- Missouri: Missouri noxious weed list (2 CSR 70-45.005).
- Ohio: Ohio endangered and threatened plants (Ohio Adm. Code 1501:18-1-03, eff. Dec. 26, 2024).
- Ohio: Ohio invasive plant species (Ohio Adm. Code 901:5-30-01).
- Oklahoma: Oklahoma Noxious Weed Law (2 O.S. § 3-220).
- Oregon: Oregon endangered and threatened plants (OAR 603-073-0070).
- Oregon: Oregon noxious weed quarantine, A and B weeds (OAR 603-052-1200).
- Vermont: Endangered and Threatened Plants of Vermont (10 V.S.A. chap. 123; list of Feb 10, 2022).
- Vermont: Vermont Quarantine List, Regulated Introduced Plant Species Rule (eff. June 16, 2026).
- USFWS, ECOS species database (ESA listing check), October 23, 2026.
- NatureServe Explorer, Ilex glabra (global and state ranks).
- Edibility and toxicity: NC State Extension; ASPCA.
- Photo: Sandy Wolkenberg via iNaturalist · CC BY 4.0 · resized.


